FAMILIENLISTE / Ceratopsidae

Marsh, 1890

 

Ornithischia Cerapoda Marginocephalia Ceratopsia Ceratopsidae

Die Ceratopsidae ("Horngesichter"), auch Ceratopidae genannt, waren eine Gruppe von vierbeinigen, pflanzenfressenden Dinosauriern aus der oberen Kreidezeit wie Triceratops und Styracosaurus. Alle Arten bewegten sich quadruped, besaßen einen papageienähnlichen Schnabel, eine Reihe von Mahlzähnen im hinteren Teil des Kiefers sowie aufwändige Hörner und Nackenschilde. Die Gruppe gliedert sich in zwei Unterfamilien, in die Centrosaurinae und die Chasmosaurinae (auch als Ceratopsinae bezeichnet).

Die Chasmosaurinae sind in der Regel charakterisiert durch lange, dreieckige Hornschilde und gut entwickelte Stirnhörner. Der auffälligste Unterschied der Chasmosaurinae zu den Centrosaurinae bestand in den Hörnern und im Nackenschild. Das Nasenhorn war wesentlich kürzer, die Hörner über den Augen waren dafür deutlich länger. Der Nackenschild war fast ebenso lang wie der Schädel und wies große Öffnungen, die so genannten Fenestrae auf. Die Chasmosaurinae lebten bis zum Ende der Kreidezeit.

Die Centrosaurinae besaßen Hörner oder nasale Knochenkämme sowie aufwändige Stacheln auf der Oberseite des Nackenschildes. Sie werden in der Regel an ihren vorstehenden nasalen Horn, untergeordneten Stirnhörnern und an dem kurzen Nackenschild mit kleinen kegelförmigen Auswüchsen erkannt. Aus vielen Funden können die Forscher ableiten, dass sich alle juvenilen Centrosaurinae sehr ähnelten. Erst im adulten Stadium änderten sie ihr Aussehen der Spezies entsprechend. Bei den Jungtieren bestanden die nasalen Hörner aus einem quer komprimierten Wulst, ihre Nackenschilde waren leicht und verletzlich. Bei den subadulten Tieren jedoch war der Nackenschild schon fast vollständig ausgeprägt. Die Centrosaurinae hatten kurze, mächtige Schnauzen und ihre Nackenschilde waren kleiner als die der Chasmosaurinae. Die Centrosaurinae starben vor ungefähr 71 Millionen Jahren während der späten Kreidezeit aus.

Ceratopsia

Ceratopsoidea
. Ferenceratops
. Zuniceratops

-Ceratopsidae

--Centrosaurinae
--. Achelousaurus
--. Brachyceratops
--. Diabloceratops
--. Machairoceratops
--. Menefeeceratops
--. Pachyrhinosaurus
--. Sinoceratops
--. Stellasaurus
--. Wendiceratops
--. Xenoceratops

---Centrosaurini
---. Centrosaurus (Typ)
---. Coronosaurus
---. Einiosaurus
---. Rubeosaurus
---. Spinops
---. Styracosaurus

---. Albertaceratopsini
---. Albertaceratops
---. Lokiceratops
---. Medusaceratops

---- Nasutoceratopsini
-----. Avaceratops
-----. Crittendenceratops
-----. Furcatoceratops
-----. Nasutoceratops
-----. Yehuecauhceratops

------Chasmosaurinae
------. Agujaceratops
------. Anchiceratops
------. Arrhinoceratops
------. Bisticeratops
------. Bravoceratops
------. Chasmosaurus (Typ)
------. Coahuilaceratops
------. Cryptarcus
------. Eotriceratops
------. Kosmoceratops
------. Mercuriceratops
------. Mojoceratops
------. Navajoceratops
------. Ojoceratops
------. Pentaceratops
------. Regaliceratops
------. Spiclypeus
------. Sierraceratops
------. ~Tatankaceratops
------. Terminocavus
------. Titanoceratops
------. Triceratops
------. Utahceratops
------. Vagaceratops

 

Weitere Informationen

Anchiceratops, a new genus of horned dinosaurs from the Edmonton Cretaceous of Alberta ; with Discussion of the origin of the ceratopsian crest and the brain casts of Anchiceratops and Trachodon / Barnum Brown, 1914 / Bulletin of the AMNH; v. 33, Article 33

A Centrosaurine (Dinosauria: Ceratopsia) from the Aguja Formation (Late Campanian) of Northern Coahuila, Mexico / Héctor E. Rivera-Sylva, Brandon P. Hedrick, Peter Dodson, 2016 / PLoS ONE 11(4): e0150529 /PDF

A chasmosaurine ceratopsid premaxilla from the basal sandstone of the Hell Creek Formation, Montana / John Scannella, 2020 / Vertebrate Anatomy Morphology Palaeontology, Volume 8/PDF

A new ceratopsian dinosaur from the Upper Cretaceous of Montana, with note on Hypacrosaurus / Charles W. Gilmore, 1914 / Smithsonian Miscellaneous Collections 63(3): 1-10

A new Ceratopsid dinosaur (Centrosaurinae: Nasutoceratopsini) from the Fort Crittenden Formation, Upper Cretaceous (Campanium) of Arizona / Sebastian Dalman, John-Paul Michael Hodnett, Asher C. Lichtig, Spencer G. Lucas, 2018 / Fossil Record 6. New Mexico Museum of Natural History and Science Bulletin 79

A new, transitional centrosaurine ceratopsid from the Upper Cretaceous Two Medicine Formation of Montana and the evolution of the ‘Styracosaurus-line' dinosaurs / John P. Wilson, Michael J. Ryan, David C. Evans, 2020 / Royal Society Open Science, 7200284PDF

A Re-Evaluation of the Chasmosaurine Ceratopsid Genus Chasmosaurus (Dinosauria: Ornithischia) from the Upper Cretaceous (Campanian) Dinosaur Park Formation of Western Canada / James A. Campbell, Michael J. Ryan, Robert B. Holmes, Claudia J. Schröder-Adams, 2016 / PLoS ONE 11(1): e0145805 /PDF

A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia / Scott D. Sampson, Eric K. Lund, Mark A. Loewen, Andrew A. Farke, Katherine E. Clayton, 2013 / Proc. R. Soc. B 7 September 2013 vol. 280 no. 1766 20131186 /PDF

A Subadult Specimen of Rubeosaurus ovatus (Dinosauria: Ceratopsidae), with Observations on Other Ceratopsids from the Two Medicine Formation / Andrew T. McDonald, 2011 / PLoS ONE 6(8): e22710 /PDF

A subadult individual of Styracosaurus albertensis (Ornithischia: Ceratopsidae) with comments on ontogeny and intraspecific variation in Styracosaurus and Centrosaurus / Caleb M. Brown, Robert B. Holmes, Philip J. Currie, 2020 / Vertebrate Anatomy Morphology Palaeontology 8:67–95 67. ISSN 2292-1389 /PDF

An Immature Pachyrhinosaurus perotorum (Dinosauria: Ceratopsidae) Nasal Reveals Unexpected Complexity of Craniofacial Ontogeny and Integument in Pachyrhinosaurus / Anthony R. Fiorillo, Ronald S. Tykoski, 2013 / PLoS ONE 8(6): e65802. doi:10.1371/journal.pone.0065802 /PDF

A New Centrosaurine Ceratopsid, Machairoceratops cronusi gen et sp. nov., from the Upper Sand Member of the Wahweap Formation (Middle Campanian), Southern Utah / Eric K. Lund, Patrick M. O’Connor, Mark A. Loewen, Zubair A. Jinnah, 2016 / PLoS ONE 11(5): e0154403 /PDF

A new ceratopsid from the Foremost Formation (middle Campanian) of Alberta / Michael J. Ryan, David C. Evans, Kieran M. Shepherd, 2012 / Canadian Journal of Earth Sciences 49 (10): 1251 – 1262

A new genus and species of Ceratopsia from New Mexico, Pentaceratops sternbergii / Charles Hazelius Sternberg, 1923 / American Museum novitates Nr. 93

A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska / Anthony R. Fiorillo, Ronald S. Tykoski, 2012 / Acta Palaeontologica Polonica 57 (3), 2012: 561-573 /PDF

A Re-Evaluation of the Chasmosaurine Ceratopsid Genus Chasmosaurus (Dinosauria: Ornithischia) from the Upper Cretaceous (Campanian) Dinosaur Park Formation of Western Canada / James A. Campbell, Michael J. Ryan, Robert B. Holmes, Claudia J. Schröder-Adams, 2016 / PLoS ONE 11(1): e0145805 /PDF

A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia / Scott D. Sampson, Eric K. Lund, Mark A. Loewen, Andrew A. Farke, Katherine E. Clayton, 2013 / Proc. R. Soc. B. 7 September 2013. Vol. 280 No. 1766 20131186 /PDF

A Spinops sternbergorum (Ornithsichia: Ceratopsia) parietal from the Dinosaur Park Formation (upper Campanian) of Muddy Lake, Saskatchewan / Jordan C. Mallon, Michael J. Ryan, Tim T. Tokaryk, 2025 / The Canadian Field-Naturalist 138(4): 294– 299 /PDF

An Immature Pachyrhinosaurus perotorum (Dinosauria: Ceratopsidae) Nasal Reveals Unexpected Complexity of Craniofacial Ontogeny and Integument in Pachyrhinosaurus / Anthony R. Fiorillo, Ronald S. Tykoski, 2013 / PLoS ONE 8(6): e65802 /PDF

Brachyceratops, a ceratopsian dinosaur from the Two Medicine formation of Montana, with notes on associated fossil reptiles / Charles W. Gilmore, 1917 / USGS Professional Paper 103

Comparative cranial osteology of subadult eucentrosauran ceratopsid dinosaurs from the Two Medicine Formation, Montana, indicates sequence of ornamentation development and complex supraorbital ontogenetic change / John P. Wilson, John B. Scannella, 2021 / Acta Palaeontologica Polonica 66 (4), 2021: 797-814 / PDF

Cranial Anatomy of Wendiceratops pinhornensis gen. et sp. nov., a Centrosaurine Ceratopsid (Dinosauria: Ornithischia) from the Oldman Formation (Campanian), Alberta, Canada, and the Evolution of Ceratopsid Nasal Ornamentation / David C. Evans, Michael J. Ryan, 2015 / PLoS ONE 10(7): e0130007 /PDF

Craniofacial ontogeny in Centrosaurus apertus / Joseph A. Frederickson​, Allison R. Tumarkin-Deratzian, 2014
PeerJ 2:e252 /PDF

Evidence of Combat in Triceratops / Andrew A. Farke, Ewan D. S. Wolff, Darren H. Tanke, 2009 / PLoS ONE 4(1): e4252 /PDF

Horn Use in Triceratops (Dinosauria: Ceratopsidae): Testing Behavioral Hypotheses Using Scale Models / Andrew A. Farke, 2010 / Palaeontologia Electronica 7 (1): 1-10. 2010

Lokiceratops rangiformis gen. et sp. nov. (Ceratopsidae: Centrosaurinae) from the Campanian Judith River Formation of Montana reveals rapid regional radiations and extreme endemism within centrosaurine dinosaurs / Mark A. Loewen​​, Joseph J. W. Sertich​, Scott Sampson, Jingmai K. O’Connor, Savhannah Carpenter, Brock Sisson, Anna Øhlenschlæger, Andrew A. Farke, Peter J. Makovicky, Nick Longrich, David C. Evans, 2024 / PeerJ 12:e17224 /PDF

Long-horned Ceratopsidae from the Foremost Formation (Campanian) of southern Alberta / Caleb M. Brown​, 2018 / PeerJ 6:e4265 /PDF

New Horned Dinosaurs from Utah Provide Evidence for Intracontinental Dinosaur Endemism / Scott D. Sampson, Mark A. Loewen, Andrew A. Farke, Eric M. Roberts, Catherine A. Forster, Joshua A. Smith, Alan L. Titus, 2010 / PLoS ONE 5(9): e12292 / PDF

New information on the holotype of “Chasmosaurus” russelli (Ornithischia: Ceratopsidae) necessitates the establishment of a new genus to receive the species / Robert B. Holmes, Jordan C. Mallon, Michael J. Ryan, David C. Evans, 2026 / Canadian Journal of Earth Sciences, Volume 63, 2026 /PDF

New information on the skull of Zuniceratops christopheri, a neoceratopsian dinosaur from the Cretaceous Moreno Hill Formation, New Mexico / Douglas G. Wolfe, 2000 / New Mexico Museum of Natural History and Science Bulletin No. 17, pp. 92-94

New insights into chasmosaurine (Dinosauria: Ceratopsidae) skulls from the Upper Cretaceous (Campanian) of Alberta, and an update on the distribution of accessory frill fenestrae in Chasmosaurinae / James A. Campbell​, Michael J. Ryan, Claudia J. Schröder-Adams, David C. Evans, Robert B. Holmes, 2018 / PeerJ 6:e5194 /PDF

Spiclypeus shipporum gen. et sp. nov., a Boldly Audacious New Chasmosaurine Ceratopsid (Dinosauria: Ornithischia) from the Judith River Formation (Upper Cretaceous: Campanian) of Montana, USA / Jordan C. Mallon, Christopher J. Ott, Peter L. Larson, Edward M. Iuliano, David C. Evans, 2016 / PLoS ONE 11(5): e0154218 /PDF

Stratigraphic Reassessment of the Mexican Chasmosaurine Coahuilaceratops magnacuerna as the First Diagnostic Dinosaur Remains from the Cerro Huerta Formation (Lower Maastrichtian) Supporting the Southern Origin of the Triceratopsini / Daniela Barrera Guevara, Belinda Espinosa Chávez, Claudia Inés Serrano Brañas, Claudio de León Dávila, Daniel Posada Martinez, Elizabeth Freedman Fowler, Denver Fowler, 2024 / Diversity, Volume 16, Issue 7, 10.3390/d16070390 /PDF

The postcranial skeleton of Vagaceratops irvinensis (Dinosauria, Ceratopsidae) / Robert B. Holmes, 2014 / Vertebrate Anatomy Morphology Palaeontology, 1, 1-21 / PDF

The skeleton of Styracosaurus with the description of a new species / Barnum Brown, Erich Maren Schlaikjer, 1937 / American Museum novitates; no. 955

Torosaurus Is Not Triceratops: Ontogeny in Chasmosaurine Ceratopsids as a Case Study in Dinosaur Taxonomy / Nicholas R. Longrich, Daniel J. Field, 2012 / PLoS ONE 7(2): e32623 /PDF

Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs: evidence from the Campanian of New Mexico / Denver W. Fowler​, Elizabeth A. Freedman Fowler, 2020 / PeerJ 8:e9251 /PDF

Turanoceratops tardabilis-the first ceratopsid dinosaur from Asia / Hans-Dieter Sues, Alexander Averianov, 2009
The Science of Nature, 06/2009; 96(5):645-52

Zuniceratops christopheri n. gen. & n. sp., a ceratopsian dinosaur from the Moreno Hill Formation (Cretaceous, Turonian) of west-central New Mexico / Douglas G. Wolfe, James I. Kirkland, 1998 / New Mexico Museum of Natural History and Science Bulletin 24: 307–317

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Bildlizenzen

Schädel des Triceratops / © Kit Dunsmore (Flickr):
Creative Commons ShareAlike 2.0 Generic (CC BY-SA 2.0)

Skelett des Chasmosaurus / © Jonathan Crowe (Flickr):
Creative Commons NonCommercial-NoDerivs 2.0 Generic (CC BY-NC-ND 2.0)

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Grafiken und Illustrationen von Raul Lunia